Towers

Three paths per school, thirty-six in all. A path fixes what the tower is for and opens ten upgrade slots — three that every tower in the game has, and seven of its own.

Every path blends three roles out of six: single-target damage, area damage, crowd control, damage over time, support (helping neighbouring towers) and debuff (weakening enemies). Slot costs below are the price of that slot's first level; each level after multiplies it.

ConflagrationPyromancy path A, starting with Burn and Burn Spread.

School: PyromancyPath: ARoles: area damage · damage over time · debuff

Abilities

On the tower from the moment it is built, before any upgrade.

Burn

Lights enemies on fire — burns for 3s at ~30% tower damage per tick.

Burn Spread

Burns leap to nearby enemies on hit, chaining the DoT through clusters.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+2.5 damage	20g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.08 shots/s	18g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.25 range	22g	×1.45	6
Splash Radius	Each shot's fire-burst covers a wider ring.	+0.3 splash radius	25g	×1.45	8
Burn Duration	Burn DoT ticks on the target for longer before fading.	+0.25 damage	20g	×1.5	6
Burn Spread	Burn jumps to more nearby enemies on contact.	+1.35 damage	40g	×1.6	4
Fire Resist Shred	Strips fire resistance so follow-up hits land harder.	+0.9 damage; level 1 adds Sear Resist	30g	×1.7	no cap
Cascade Chance	Higher roll for the burn to cascade onto neighbours on contact.	+5 to the ability; level 1 adds Fire Cascade	35g	×1.55	10
AoE Damage %	Each level widens the burn-spread AoE so more enemies catch the splash.	+0.2 splash radius	28g	×1.45	8
Ignition Aura	Wider ignition aura — more allied towers catch the burn-spread buff.	+0.3 aura range; level 1 adds Shared Ignition	40g	×1.6	5

Its schoolPyromancy→Path BPyroclasm→Path CEmberheart→

PyroclasmPyromancy path B, starting with Burn and Melt Focus.

School: PyromancyPath: BRoles: single-target damage · damage over time · crowd control

Abilities

On the tower from the moment it is built, before any upgrade.

Burn

Lights enemies on fire — burns for 3s at ~30% tower damage per tick.

Melt Focus

Stacks a burn-damage amplifier on the target, up to ~8 stacks per hit.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	22g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.1 shots/s	20g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	24g	×1.45	6
Burn Intensity	Each burn stack ticks harder on a single target.	+0.05 damage	22g	×1.5	no cap
Max Stacks	Cap on simultaneous burn stacks lifts — more compound damage on tough targets.	+1 to the ability	50g	×1.8	6
Root Duration	Slag-chained enemies stay rooted for longer.	+0.3 s control; level 1 adds Slag Chains	35g	×1.55	5
Root Chance	Higher roll that the burn chains into a slag root.	+3% proc chance; level 1 adds Slag Chains	30g	×1.5	8
Magma Pool Size	Magma pools left on enemy death cover more ground.	+0.2 splash radius; level 1 adds Molten Collapse	40g	×1.55	6
Magma Pool Duration	Magma pools linger longer after spawning.	+0.5 damage; level 1 adds Molten Collapse	35g	×1.5	5
Crit Chance	Higher chance for a shot to roll a critical hit.	+0.27 damage	25g	×1.55	10

Its schoolPyromancy→Path AConflagration→Path CEmberheart→

EmberheartPyromancy path C, starting with Burn and Aura Attack Speed 8.

School: PyromancyPath: CRoles: support · area damage · damage over time

Abilities

On the tower from the moment it is built, before any upgrade.

Burn

Lights enemies on fire — burns for 3s at ~30% tower damage per tick.

Aura Attack Speed 8

Carried by every tower on this path.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+1.5 damage	18g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.06 shots/s	16g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.2 range	20g	×1.45	6
Aura AS Strength	Allied towers in aura fire faster.	+2% aura strength	30g	×1.55	no cap
Aura Range	Aura covers more cells around the tower.	+0.3 aura range	35g	×1.5	6
Shared Ignition	Boosts every aura — allied towers near this one fire faster, hit harder, and reach further.	+2% aura strength; level 1 adds Shared Ignition	40g	×1.6	8
Phoenix Surge Dmg	Phoenix Surge burn ticks harder on the target.	+3 to the ability; level 1 adds Phoenix Surge	45g	×1.55	no cap
Surge Frequency	Phoenix Surge ticks more often (interval shortens, floor ~0.2s).	+0.05 to the ability; level 1 adds Phoenix Surge	50g	×1.65	6
Phoenix Wing	Phoenix Surge burn lasts longer before falling off.	+0.5 to the ability; level 1 adds Phoenix Surge	35g	×1.5	5
Buff Duration	Every status the tower applies sticks on the target for longer.	+0.5 s control	25g	×1.45	6

Its schoolPyromancy→Path AConflagration→Path BPyroclasm→

PermafrostCryomancy path A, starting with Slow and Frost Patch.

School: CryomancyPath: ARoles: crowd control · area damage · debuff

Abilities

On the tower from the moment it is built, before any upgrade.

Slow

Slows movement by 25% for 2.5 seconds.

Frost Patch

Hits chill the ground briefly, slowing enemies that pass through.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+1.5 damage	18g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.07 shots/s	16g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.25 range	22g	×1.45	6
Slow Strength	Slow stacks bite deeper — enemies crawl slower.	+3% slow	20g	×1.5	10
Frost Patch Size	Frost patches dropped underfoot cover more ground.	+0.2 splash radius	28g	×1.5	8
Patch Duration	Frost patches linger longer after being laid.	+0.5 damage	25g	×1.45	no cap
Armor Shred	Each hit strips more armor — frost shots bite through plate.	+0.6 damage; level 1 adds Armor Crack	30g	×1.55	no cap
Freeze Chance	Higher roll that a hit fully freezes the target.	+2% proc chance; level 1 adds Deep Freeze	40g	×1.65	10
Freeze Duration	Frozen enemies stay locked for longer.	+0.3 s control	45g	×1.6	5
Dmg Amp on Frozen	Each level adds raw damage to every shot — a flat hit-power bump.	+0.72 damage	35g	×1.55	8

Its schoolCryomancy→Path BShatter→Path CFrostweave→

ShatterCryomancy path B, starting with Slow and Brittle Strike.

School: CryomancyPath: BRoles: single-target damage · crowd control · debuff

Abilities

On the tower from the moment it is built, before any upgrade.

Slow

Slows movement by 25% for 2.5 seconds.

Brittle Strike

Target takes 25% more damage from any source for 3 seconds.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+2.5 damage	20g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.08 shots/s	18g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.25 range	22g	×1.45	6
Brittle Bonus	Each hit lays a brittle damage-amp on the target — follow-up shots land harder.	+4 to the ability	22g	×1.5	no cap
Freeze Chance	Higher roll that a hit fully freezes the target.	+2.5% proc chance; level 1 adds Flash Freeze	35g	×1.6	10
Shatter Damage	Shatter damage-amp on the target ramps higher per hit — every Shatter shot stacks deeper.	+3 to the ability; level 1 adds Death Shatter	45g	×1.65	8
Shatter Radius	Shatter explosion covers more cells when a frozen target dies.	+0.3 splash radius	35g	×1.5	6
Chain Freeze	Freeze jumps to more nearby enemies.	+2.4 damage	50g	×1.7	3
Crit Damage	Crit hits hit even harder.	+0.27 damage	30g	×1.55	8
Slow Duration	Slows stick to the target for longer.	+0.3 s slow	20g	×1.45	6

Its schoolCryomancy→Path APermafrost→Path CFrostweave→

FrostweaveCryomancy path C, starting with Aura Slow 10 and Frostbite.

School: CryomancyPath: CRoles: support · crowd control · damage over time

Abilities

On the tower from the moment it is built, before any upgrade.

Aura Slow 10

Carried by every tower on this path.

Frostbite

Frostbite ticks for ~20% tower damage per second for 4 seconds.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+1 damage	16g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.06 shots/s	14g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.2 range	18g	×1.45	6
Aura Slow	Allied slow aura bites deeper around the tower.	+2% aura strength	25g	×1.5	no cap
Frostbite Stacks	Frostbite stack cap rises — more compound chill damage.	+1 to the ability; level 1 adds Frostbite Stack	30g	×1.55	10
Frostbite Damage	Each frostbite stack ticks harder.	+0.001 damage; level 1 adds Frostbite Stack	28g	×1.5	no cap
Ally Crit Bonus	Allied towers in aura deal more damage.	+2% aura strength; level 1 adds Heart of Winter	40g	×1.6	8
Aura Range	Aura covers more cells around the tower.	+0.3 aura range	30g	×1.5	6
Slow Persist	Slow lingers on the target for longer after leaving a patch.	+0.5 s slow	22g	×1.45	5
Frost Armor	Frost coating slows enemies harder while it lingers.	+2% slow	35g	×1.55	6

Its schoolCryomancy→Path APermafrost→Path BShatter→

Storm ChainVoltomancy path A, starting with Chain Lightning.

School: VoltomancyPath: ARoles: area damage · single-target damage · debuff

Abilities

On the tower from the moment it is built, before any upgrade.

Chain Lightning

Each shot bounces to 2 additional targets.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Chain Targets	Chain lightning hops to more enemies.	+1.2 damage	25g	×1.5	no cap
Chain Decay	Each level increases chain-lightning damage — every bolt in the chain lands harder.	+1.2 damage	30g	×1.55	8
Shock Duration	Shocked enemies stay slowed for longer.	+0.3 s slow; level 1 adds Galvanic Shock	22g	×1.45	6
Shock Strength	Shock slow bites deeper.	+3% slow	20g	×1.5	8
Thunderclap AoE	Thunderclap stun explosion covers more cells.	+0.2 splash radius; level 1 adds Thunderclap	40g	×1.6	5
Amplify Chains	Chain bolts hit harder on every target in the chain.	+1.2 damage	35g	×1.55	no cap
Crit Chance	Higher chance for a shot to roll a critical hit.	+0.15 damage	28g	×1.55	10

Its schoolVoltomancy→Path BOvercharge→Path CGalvanic→

OverchargeVoltomancy path B, starting with Static Buildup.

School: VoltomancyPath: BRoles: single-target damage · damage over time · crowd control

Abilities

On the tower from the moment it is built, before any upgrade.

Static Buildup

Stacks static charge on hits, priming enemies for follow-up shocks.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Charge Rate	Tower fires faster as its charge builds.	+0.025 shots/s	30g	×1.55	4
Burst Damage	Burst shots land with critical-grade extra damage.	+0.9 damage	35g	×1.6	no cap
Stun Duration	Electrocution stuns hold for longer.	+0.2 s control	40g	×1.55	6
Pulse Radius	Charge-pulse AoE covers more cells on release.	+0.3 splash radius; level 1 adds Thunder Pulse	32g	×1.5	6
Lightning Rod	Allied towers in aura fire faster.	+2% aura strength; level 1 adds Lightning Rod	50g	×1.7	4
Charge Bonus Dmg	Fully-charged shots land with extra raw damage per level.	+0.72 damage	25g	×1.5	no cap
Crit Damage	Crit hits hit even harder.	+0.135 damage	28g	×1.55	8

Its schoolVoltomancy→Path AStorm Chain→Path CGalvanic→

GalvanicVoltomancy path C, starting with Aura Attack Speed 6 and Conductivity.

School: VoltomancyPath: CRoles: support · area damage · debuff

Abilities

On the tower from the moment it is built, before any upgrade.

Aura Attack Speed 6

Carried by every tower on this path.

Conductivity

Builds up to ~10 charge stacks; each adds 5% lightning damage taken.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Aura AS	Allied towers in aura fire faster.	+2% aura strength	28g	×1.55	no cap
Conductivity Stacks	Conductivity stack cap rises so the bonus chain-chance compounds further.	+1 to the ability	30g	×1.5	10
Induced Current	Allied towers in aura deal more damage (aura scales with this slot).	+2% aura strength; level 1 adds Induced Current	40g	×1.6	8
Aura Range	Aura covers more cells around the tower.	+0.3 aura range	30g	×1.5	6
Mega Bolt Damage	Perpetual Storm's mega-bolts inflict heavier burn per tick.	+2 to the ability; level 1 adds Perpetual Storm	45g	×1.65	no cap
Bolt Frequency	Perpetual Storm bolts hit more often (tick interval shrinks, floor 0.2s).	+0.05 to the ability	50g	×1.6	5
Storm Dmg/s	Passive storm damage-per-second on the target climbs.	+0.5 damage	25g	×1.5	no cap

Its schoolVoltomancy→Path AStorm Chain→Path BOvercharge→

PlagueToxicology path A, starting with Poison, Poison Cloud and Death Spore.

School: ToxicologyPath: ARoles: area damage · damage over time · debuff

Abilities

On the tower from the moment it is built, before any upgrade.

Poison

Poisons the target for 4s, ticking ~40% tower damage over the duration.

Poison Cloud

Spreads a toxic cloud that poisons everyone passing through.

Death Spore

Killed enemies release a spore cloud that poisons their allies.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Cloud Radius	Poison clouds cover more ground.	+0.2 splash radius	25g	×1.5	8
Cloud Duration	Poison clouds linger longer after spawning.	+0.5 damage	22g	×1.45	no cap
Poison Stacks	Poison stack cap rises — more compound damage on tough targets.	+1 to the ability; level 1 adds Stacking Poison	30g	×1.55	10
Tick Acceleration	Poison ticks more often (interval shrinks, floor 0.2s).	+0.05 to the ability; level 1 adds Stacking Poison	35g	×1.6	6
Death Spread	Poison jumps to more nearby enemies on death.	+1.35 damage	40g	×1.6	4
Anti-Heal	Anti-heal debuff stays on the target longer.	+0.5 to the ability; level 1 adds Anti-Heal	30g	×1.5	8
Cloud Merge	Poison clouds merge into wider patches across the field.	+0.1 splash radius	45g	×1.65	6

Its schoolToxicology→Path BVenom→Path CToxicant→

VenomToxicology path B, starting with Strong Poison and Neurotoxin.

School: ToxicologyPath: BRoles: single-target damage · damage over time · crowd control

Abilities

On the tower from the moment it is built, before any upgrade.

Strong Poison

Stronger poison: ~80% tower damage over 5 seconds.

Neurotoxin

Poisons also slow movement by 20% for 3 seconds.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Venom Potency	Venom DoT ticks harder on the target.	+0.25 damage	22g	×1.5	no cap
Neurotoxin Slow	Neurotoxin slow bites deeper.	+3% slow	20g	×1.45	10
Armor Shred/Stack	Each poison stack on the target strips more armor.	+0.3 damage; level 1 adds Acid Shred	30g	×1.55	no cap
Convulsion Chance	Higher roll for a poison tick to stun the target.	+2% proc chance	35g	×1.55	8
Accelerating DoT	Each level ramps the accelerating poison's per-tick damage higher.	+3 to the ability; level 1 adds Accelerating Poison	40g	×1.6	no cap
Toxic Aura Range	Toxic aura reaches more cells around the tower.	+0.2 aura range; level 1 adds Toxic Aura	35g	×1.55	5
Execute Threshold	Lethal-dose execute fires on enemies at higher HP %.	+1 damage	50g	×1.7	5

Its schoolToxicology→Path APlague→Path CToxicant→

ToxicantToxicology path C, starting with Poison and Aura Damage Amp 10.

School: ToxicologyPath: CRoles: support · debuff · area damage

Abilities

On the tower from the moment it is built, before any upgrade.

Poison

Poisons the target for 4s, ticking ~40% tower damage over the duration.

Aura Damage Amp 10

Carried by every tower on this path.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Damage Amp	Every poison hit chunks harder on the target.	+1.35 damage	30g	×1.55	no cap
Buff Strip	Anti-heal pulses bite deeper, suppressing enemy regen for longer.	+18 damage; level 1 adds Buff Strip	35g	×1.55	8
Resist Shred	Strips more magic resist on every hit.	+0.6 damage; level 1 adds Resist Shred	30g	×1.5	no cap
Debuff Amplifier	Each poison debuff amp stack lands harder on the target.	+3 to the ability; level 1 adds Debuff Amplifier	45g	×1.65	no cap
Wave Radius	Toxin wave covers more ground per cast.	+0.3 splash radius	28g	×1.5	6
Wave Frequency	Tower fires faster — poison waves trigger more often.	+0.05 shots/s	40g	×1.55	5
Catalytic Burst	Catalytic burst AoE covers a wider ring on detonation.	+0.2 splash radius	50g	×1.6	no cap

Its schoolToxicology→Path APlague→Path BVenom→

QuakeGeomancy path A, starting with Stunning Strike and Ground Shock.

School: GeomancyPath: ARoles: area damage · crowd control · debuff

Abilities

On the tower from the moment it is built, before any upgrade.

Stunning Strike

10% chance to stun the target for ~0.8 seconds.

Ground Shock

Every hit shakes the ground, briefly stunning the target (~0.5s).

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+4.8 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Stun Chance	Higher roll that a hit stuns the target.	+2% proc chance	25g	×1.5	10
AoE Radius	Quake shockwave covers more cells.	+0.2 splash radius	28g	×1.5	8
Rubble Slow	Rubble slows targets harder while in the patch.	+3% slow; level 1 adds Rubble Zone	22g	×1.45	8
Rubble Duration	Rubble persists longer after a quake.	+1 damage	25g	×1.5	no cap
Armor Shred	Each hit strips more armor — quake fractures shake plates loose.	+0.6 damage	30g	×1.55	no cap
Cataclysm Dmg	Cataclysm stuns hold for longer.	+0.3 to the ability; level 1 adds Cataclysm	45g	×1.6	6
Cataclysm Freq	Higher roll for the quake to escalate into a cataclysm stun.	+3 to the ability	50g	×1.65	6

Its schoolGeomancy→Path BRampart→Path CGolem→

RampartGeomancy path B, starting with Stunning Strike and Stone Barrier.

School: GeomancyPath: BRoles: support · crowd control · single-target damage

Abilities

On the tower from the moment it is built, before any upgrade.

Stunning Strike

10% chance to stun the target for ~0.8 seconds.

Stone Barrier

Drops a barrier that slows passing enemies by ~20% for 4 seconds.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+4.8 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Barrier HP	Stone-barrier slow lingers on each enemy for longer.	+0.5 to the ability	30g	×1.55	no cap
Barrier Regen	Stone-barrier slow bites deeper — adds %-points to the slow strength.	+1 to the ability	35g	×1.55	6
Ally Damage Buff	Allied towers near a barrier deal more damage.	+3% aura strength; level 1 adds Fortified Position	30g	×1.5	no cap
Ally AS Buff	Allied towers near a barrier fire faster.	+2% aura strength	28g	×1.5	no cap
Explode Damage	Barriers detonate harder when they break.	+5 to the ability; level 1 adds Barrier Explosion	40g	×1.6	no cap
Barrier Count	More barriers stand at once.	+2.4 damage	60g	×1.8	2
Counter Stun	Higher on-hit stun roll — the rampart hits back.	+1% proc chance	25g	×1.5	8

Its schoolGeomancy→Path AQuake→Path CGolem→

GolemGeomancy path C, starting with Stunning Strike and Stone Fist.

School: GeomancyPath: CRoles: single-target damage · crowd control · damage over time

Abilities

On the tower from the moment it is built, before any upgrade.

Stunning Strike

10% chance to stun the target for ~0.8 seconds.

Stone Fist

Heavy impact — each hit stuns the target for 1 second.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+4.8 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Fist Damage	Golem fist stuns hold the target for longer.	+0.3 to the ability	30g	×1.55	6
Fist Frequency	Golem fist stuns hold even longer — extra duration on top of Fist Damage.	+0.3 to the ability	35g	×1.55	6
Bleed Damage	Each bleed tick hurts more on the target.	+0.5 damage; level 1 adds Crushing Wound	25g	×1.5	no cap
Bleed Duration	Bleed lingers on the target for longer.	+0.25 damage	22g	×1.45	6
Golem HP	Golem-bleed lingers on the target for longer.	+0.3 to the ability; level 1 adds Stone Golem	40g	×1.6	no cap
Golem Damage	Each level adds raw strength to the golem's BLEED tick — the summoned bruiser hits harder.	+0.2 to the ability	35g	×1.55	no cap
Golem Regen	Each golem-bleed tick hits harder (adds raw strength per tick).	+0.5 to the ability	30g	×1.5	6

Its schoolGeomancy→Path AQuake→Path BRampart→

ReaperUmbramancy path A, starting with Execute and Death Mark.

School: UmbramancyPath: ARoles: single-target damage · debuff · damage over time

Abilities

On the tower from the moment it is built, before any upgrade.

Execute

Below 30% HP: deals massive bonus damage to finish the target.

Death Mark

Marks enemies — each stack adds ~8% damage taken, up to ~5 stacks.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3.6 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Death Mark Stacks	More death-mark stacks pile onto a single target before capping.	+1 to the ability	30g	×1.55	5
Execute Threshold	Execute fires on targets at higher HP %.	+1.5 damage; level 1 adds Final Harvest	40g	×1.65	8
Soul Bleed	Soul-bleed DoT chews more % maxHP per second.	+0.25 damage; level 1 adds Soul Bleed	35g	×1.6	no cap
Mark Dmg Amp	Marked targets take amplified damage on every follow-up hit.	+1.44 damage	28g	×1.5	no cap
Reap Speed	Each kill refunds part of the tower's next-shot cooldown.	+25% cooldown back on a kill	45g	×1.65	4
Kill Chain	Each kill spills damage onto nearby enemies, chaining harder per level.	+1.44 damage	50g	×1.7	no cap
Crit Chance	Higher chance for a shot to roll a critical hit.	+0.36 damage	25g	×1.5	10

Its schoolUmbramancy→Path BPhantom→Path CUmbral→

PhantomUmbramancy path B, starting with Fear.

School: UmbramancyPath: BRoles: area damage · crowd control · debuff

Abilities

On the tower from the moment it is built, before any upgrade.

Fear

20% chance to terrify the target — they flee backward for 1.5 seconds.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3.6 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Fear Chance	Higher roll that a hit fears the target into running.	+3% proc chance	30g	×1.55	10
Fear Duration	Feared enemies stay fleeing for longer.	+0.2 s control	35g	×1.55	6
Fear Spread	Fear jumps to more nearby enemies.	+1.8 damage; level 1 adds Mass Hysteria	45g	×1.65	3
Resist Shred	Strips more magic resist on every hit.	+0.6 damage; level 1 adds Resist Shred	28g	×1.5	no cap
Dread Stacks	Haunted-fear lingers on the target for longer.	+0.3 to the ability; level 1 adds Haunted Ground	30g	×1.5	6
Phantom Duration	Phantom shroud holds fear/stun on the target for longer.	+0.2 s control	35g	×1.55	6
Dmg Reduction	Higher chance for hits to interrupt the target with crowd-control.	+1.5% proc chance	25g	×1.5	8

Its schoolUmbramancy→Path AReaper→Path CUmbral→

UmbralUmbramancy path C, starting with Shadow Cloak and Blind.

School: UmbramancyPath: CRoles: support · single-target damage · crowd control

Abilities

On the tower from the moment it is built, before any upgrade.

Shadow Cloak

Carried by every tower on this path.

Blind

Blinds the target — their hit chance drops 30% for 2 seconds.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3.6 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Blind Strength	Blind miss-chance bites deeper — enemies whiff more often.	+3 to the ability	25g	×1.5	8
Blind Duration	Blind sticks on the target for longer.	+0.3 s control	22g	×1.45	6
Ambush Damage	Every shot lays an ambush damage-amp — follow-up hits on the target land harder.	+5 to the ability; level 1 adds Ambush Pact	35g	×1.55	no cap
Banish Damage	Banish stuns hold for longer.	+0.3 to the ability; level 1 adds Dimensional Rift	45g	×1.65	8
Banish Frequency	Higher roll for a hit to banish the target.	+3 to the ability	50g	×1.65	5
Phase Speed	Phased tower fires faster.	+0.15 shots/s	35g	×1.55	6
Cloak Range	Shadowcloak aura covers more cells.	+0.3 aura range	30g	×1.5	5

Its schoolUmbramancy→Path AReaper→Path BPhantom→

RadianceRadiance path A, starting with Reveal, Solar Flare and Mending Light.

School: RadiancePath: ARoles: area damage · support · damage over time

Abilities

On the tower from the moment it is built, before any upgrade.

Reveal

Sees through stealth — strips invisibility from any enemy in range.

Solar Flare

Wide solar burst sears all enemies in range for ~40% damage over 3s.

Mending Light

Aura: heals nearby allied towers / repairs barriers.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3.3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Flash Radius	Solar Flare AoE covers more cells.	+0.2 splash radius	25g	×1.5	8
Sunbeam Focus	Solar Flare burn ticks harder on the target.	+1 to the ability	30g	×1.55	6
Mark of Light	Marked targets take amplified damage on every follow-up hit.	+2 to the ability; level 1 adds Mark of Light	35g	×1.55	no cap
Daybreak Dmg	Daybreak burst burn ticks harder.	+3 to the ability; level 1 adds Daybreak	45g	×1.65	no cap
Daybreak Freq	Daybreak burn lingers on the target for longer.	+0.4 to the ability	50g	×1.6	5
Aegis Burn	Strips magic resistance with each Aegis tick.	+1 to the ability; level 1 adds Aegis Burn	40g	×1.6	6
Sight Beyond	Stretches the tower's targeting reach further out.	+0.3 range	20g	×1.45	6

Its schoolRadiance→Path BJudgment→Path CSanctuary→

JudgmentRadiance path B, starting with Reveal and Holy Smite.

School: RadiancePath: BRoles: single-target damage · debuff · crowd control

Abilities

On the tower from the moment it is built, before any upgrade.

Reveal

Sees through stealth — strips invisibility from any enemy in range.

Holy Smite

Smites enemies with a 10% damage-amp debuff for 3 seconds.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3.3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Smite Bonus	Smite-weakness debuff bites deeper on the target.	+2 to the ability	25g	×1.5	no cap
Purge Stun	Purge stuns hold for longer.	+0.2 s control; level 1 adds Purifying Light	35g	×1.55	5
Sentence Dmg Amp	Judgment-sentenced enemies take amplified damage.	+3 to the ability; level 1 adds Divine Sentence	40g	×1.6	no cap
Sentence Duration	Judgment debuff sticks on the target for longer.	+1 to the ability	35g	×1.55	6
Sentence Resist Shred	Sentenced enemies lose more magic resistance.	+0.9 damage	38g	×1.55	8
Own Dmg vs Judged	Each shot lands harder on enemies under the tower's judgment.	+1.32 damage	30g	×1.5	no cap
Crit Chance	Higher chance for a shot to roll a critical hit.	+0.33 damage	25g	×1.55	10

Its schoolRadiance→Path ARadiance→Path CSanctuary→

SanctuaryRadiance path C, starting with Reveal, Aura Slow 15 and Aura Range 10.

School: RadiancePath: CRoles: support · crowd control · area damage

Abilities

On the tower from the moment it is built, before any upgrade.

Reveal

Sees through stealth — strips invisibility from any enemy in range.

Aura Slow 15

Carried by every tower on this path.

Aura Range 10

Carried by every tower on this path.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3.3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Zone Slow	Sanctuary zone slows enemies harder.	+2% slow	22g	×1.5	no cap
Zone Damage	Sanctuary zone burns enemies harder per second.	+0.5 damage	28g	×1.5	no cap
Ally Dmg Buff	Allied towers in the zone deal more damage.	+2% aura strength	30g	×1.55	no cap
Ally Range Buff	Allied towers in the zone gain extra targeting range.	+0.15% aura strength	28g	×1.5	6
Ally AS Buff	Allied towers in the zone fire faster.	+1.5% aura strength	28g	×1.5	no cap
Blind Chance	Higher roll for a hit to blind the target.	+2 to the ability; level 1 adds Holy Blind	35g	×1.55	8
Pulse Stun	Absolution pulse stuns hold for longer.	+0.2 s control; level 1 adds Absolution Pulse	45g	×1.6	5

Its schoolRadiance→Path ARadiance→Path BJudgment→

DisintegrationArcanum path A, starting with Adaptive Damage and Arcane Erosion.

School: ArcanumPath: ARoles: single-target damage · debuff · area damage

Abilities

On the tower from the moment it is built, before any upgrade.

Adaptive Damage

Each consecutive hit on the same target adds +10% damage (resets on switch).

Arcane Erosion

Stacks magical erosion (up to ~10) — each strips 3% magic resist.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Erosion Stacks	Erosion stack cap rises — compound resist-shred sticks deeper.	+1 to the ability	28g	×1.5	10
Erosion Strength	Each erosion stack strips more magic resistance.	+0.15 damage	30g	×1.55	no cap
Beam Ramp	Sustained beam on a single target ramps up damage faster.	+2 to the ability; level 1 adds Focused Beam	35g	×1.55	no cap
Pierce Count	Arcane beam pierces more enemies in a line.	+2 damage	45g	×1.65	3
Void Zone Dmg	Void zones strip more magic resistance from enemies inside.	+3 to the ability; level 1 adds Nullification	40g	×1.6	no cap
Void Zone Duration	Void zones linger longer on the field.	+0.5 damage	35g	×1.55	5
%HP Damage	Beam-amplification holds on the target for longer.	+0.3 to the ability; level 1 adds Focused Beam	50g	×1.7	no cap

Its schoolArcanum→Path BSpellweave→Path CSiphon→

SpellweaveArcanum path B, starting with Rune Placement.

School: ArcanumPath: BRoles: area damage · support · crowd control

Abilities

On the tower from the moment it is built, before any upgrade.

Rune Placement

Plants arcane runes that detonate periodically for AoE damage.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Max Runes	More runes can be placed and active at once.	+1.5 damage	30g	×1.55	6
Rune Damage	Every shot deals more damage — the tower channels rune power into hits.	+1.2 damage	25g	×1.5	no cap
Rune Cascade	Rune-cascade burn ticks harder on the target.	+3 to the ability; level 1 adds Rune Link	40g	×1.65	3
Tripwire Dmg	Tripwire shots hit harder on every triggered hit.	+1.2 damage	30g	×1.55	no cap
Overload Dmg	Overload rune burst burns enemies harder per level.	+0.3 to the ability; level 1 adds Living Spellwork	50g	×1.65	no cap
Rune Regen	Tower fires faster as runes regenerate between casts.	+0.02 shots/s	35g	×1.55	6
Ally Buff on Rune	Allied towers near a rune deal more damage.	+2% aura strength	28g	×1.5	8

Its schoolArcanum→Path ADisintegration→Path CSiphon→

SiphonArcanum path C, starting with Arcane Siphon.

School: ArcanumPath: CRoles: support · damage over time · debuff

Abilities

On the tower from the moment it is built, before any upgrade.

Arcane Siphon

Drains stats from the target — each stack weakens them further.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Siphon Rate	Each hit siphons more enemy strength into a weakness debuff.	+1 to the ability	25g	×1.5	no cap
Redistribute	Allied towers in aura fire faster.	+2% aura strength; level 1 adds Essence Redistribute	35g	×1.55	8
Mana Burn	Mana-burn DoT chews more % maxHP per second.	+0.15 damage	30g	×1.55	no cap
Wellspring Scaling	Wellspring amplifies damage on the target further.	+0.5 to the ability; level 1 adds Arcane Wellspring	40g	×1.6	no cap
Husk Threshold	Husk-burn lingers on the target for longer.	+0.5 to the ability	45g	×1.65	5
Husk Burst	Husk-burn ticks harder on the target.	+1.5 to the ability; level 1 adds Husk Drain	35g	×1.55	no cap
Aura Range	Wellspring aura covers more cells around the tower.	+0.3 aura range	30g	×1.5	6

Its schoolArcanum→Path ADisintegration→Path BSpellweave→

CycloneAeromancy path A, starting with Knockback and Dust Devil.

School: AeromancyPath: ARoles: area damage · crowd control · damage over time

Abilities

On the tower from the moment it is built, before any upgrade.

Knockback

Knocks the target back ~0.5 tiles on hit.

Dust Devil

Whirlwind on hit slows nearby enemies by ~20% for 3 seconds.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Tornado Duration	Tornadoes linger longer on the field.	+0.5 damage	25g	×1.5	no cap
Tornado Count	More tornadoes can be active at once.	+1.2 damage	50g	×1.75	3
Windshear Dmg	Windshear DoT chews more % maxHP per second.	+0.1 damage; level 1 adds Windshear	30g	×1.55	no cap
Mega Cyclone Stun	Merged mega-cyclone stuns hold for longer.	+0.3 s control; level 1 adds Mega Cyclone	40g	×1.6	6
Pull Strength	Tornado pull slows enemies caught inside harder.	+0.2 to the ability	28g	×1.5	6
Tornado Size	Tornadoes cover more cells.	+0.2 splash radius	30g	×1.55	8
Growth Rate	Tornado slow lingers on the target for longer.	+0.3 to the ability	35g	×1.55	8

Its schoolAeromancy→Path BZephyr→Path CShrike→

ZephyrAeromancy path B, starting with Aura Range 15 and Aura Slow 10.

School: AeromancyPath: BRoles: support · crowd control · debuff

Abilities

On the tower from the moment it is built, before any upgrade.

Aura Range 15

Carried by every tower on this path.

Aura Slow 10

Carried by every tower on this path.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Headwind Slow	Headwind slows enemies harder while in the gust.	+2% slow	22g	×1.5	no cap
Tailwind Range	Allied towers in tailwind aura gain extra targeting range.	+0.15% aura strength	28g	×1.5	6
Ally AS Buff	Allied towers in wind aura fire faster.	+2% aura strength	25g	×1.5	no cap
Ground Flyers	Grounded flyers slow harder once forced down.	+5 to the ability; level 1 adds Ground Flyers	40g	×1.6	4
Wind Reversal	Wind reversal fear holds enemies fleeing for longer when it triggers.	+1 to the ability; level 1 adds Wind Reversal	50g	×1.65	5
Reversal Freq	Higher roll that a hit reverses the enemy's direction.	+3 to the ability	45g	×1.6	5
Buff Strip	Every shot lands with extra damage from stripped enemy buffs.	+0.96 damage	35g	×1.55	4

Its schoolAeromancy→Path ACyclone→Path CShrike→

ShrikeAeromancy path C, starting with Knockback and Bleed.

School: AeromancyPath: CRoles: single-target damage · crowd control · area damage

Abilities

On the tower from the moment it is built, before any upgrade.

Knockback

Knocks the target back ~1 tile on hit.

Bleed

Wind-shorn cuts bleed for ~25% tower damage over 3 seconds.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Knockback Force	Each hit knocks targets back further.	+0.2 knockback	25g	×1.5	8
Bleed Damage	Each bleed tick hurts more on the target.	+0.25 damage	22g	×1.5	no cap
Ricochet Count	Shots ricochet to more enemies.	+1.6 damage; level 1 adds Ricochet	35g	×1.6	4
Collision Dmg	Collision stuns hold for longer.	+0.3 to the ability; level 1 adds Collision Damage	40g	×1.6	no cap
Gale Lance	Gale Lance knocks targets back further.	+0.5 to the ability; level 1 adds Gale Lance	45g	×1.65	no cap
Lance Frequency	Gale Lance knocks targets back further (stacks with Gale Lance).	+0.3 to the ability	40g	×1.55	6
Bleed Stacks	Bleed stack cap rises — more compound bleed damage on tough targets.	+1 to the ability	30g	×1.5	10

Its schoolAeromancy→Path ACyclone→Path BZephyr→

OvergrowthDruidism path A, starting with Thorns and Briar Patch.

School: DruidismPath: ARoles: area damage · damage over time · crowd control

Abilities

On the tower from the moment it is built, before any upgrade.

Thorns

Vines deal ~20% tower damage per second for 3 seconds.

Briar Patch

Drops briars at the target — passing enemies are slowed 15% for 3s.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Vine Damage	Each thorn tick hurts more on the target.	+0.25 damage	22g	×1.5	no cap
Patch Coverage	Thorn patches cover more cells.	+0.2 splash radius	28g	×1.5	8
Root Chance	Higher roll that a hit roots the target in place.	+3% proc chance; level 1 adds Entangle	30g	×1.55	10
Root Duration	Rooted enemies stay locked for longer.	+0.2 s control	28g	×1.5	6
Auto Spread	Auto-spread roots hold for longer (stacks with Root Duration).	+0.3 to the ability; level 1 adds Living Labyrinth	40g	×1.6	no cap
Seed Pod Freq	Seed pods tick more often (interval shrinks, floor 0.2s).	+0.1 to the ability; level 1 adds Seed Pod	35g	×1.55	6
Perma-Root Count	Higher chance for hits to trigger a permanent root.	+0.5% proc chance	50g	×1.7	3

Its schoolDruidism→Path BDruid→Path CBeastcaller→

DruidDruidism path B, starting with Thorn Aura and Natural Decay.

School: DruidismPath: BRoles: support · area damage · debuff

Abilities

On the tower from the moment it is built, before any upgrade.

Thorn Aura

Aura: damaged allies retaliate with a thorn DoT against the attacker.

Natural Decay

Each hit accelerates decay — strips 5 armor per hit.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Aura Thorn Dmg	Allied towers in aura thorn-tick enemies harder.	+0.15 damage	25g	×1.5	no cap
Natural Decay	Strips more armor on every hit.	+0.3 damage	28g	×1.5	no cap
Summon Buff %	Allied towers in aura deal more damage.	+3% aura strength; level 1 adds Summon Buff	30g	×1.55	no cap
Wilting Pulse	Wilting-pulse anti-heal lingers on the target longer.	+0.5 to the ability; level 1 adds Healing Burst	35g	×1.55	8
Season Cycle Speed	Season cycle damage-amp grows stronger per level.	+0.5 to the ability; level 1 adds Cycle of Seasons	40g	×1.6	5
Season Strength	All season-aura buffs scale stronger on allies.	+1.8% aura strength	35g	×1.55	no cap
Aura Range	Nature aura covers more cells around the tower.	+0.3 aura range	28g	×1.5	6

Its schoolDruidism→Path AOvergrowth→Path CBeastcaller→

BeastcallerDruidism path C, starting with Wolf Summon.

School: DruidismPath: CRoles: single-target damage · support · crowd control

Abilities

On the tower from the moment it is built, before any upgrade.

Wolf Summon

Summons a wolf that lunges into melee and rakes enemies.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Wolf Damage	Wolf bleed ticks harder on the target.	+0.2 to the ability	25g	×1.5	no cap
Wolf HP	Wolf bleed lingers on the target for longer.	+0.15 to the ability	22g	×1.5	no cap
Hawk Marks	Hawks mark more enemies at once.	+1.35 damage; level 1 adds Hawk Summon	40g	×1.6	4
Mark Dmg Amp	Marked targets take amplified damage on every follow-up hit.	+1.44 damage	30g	×1.55	no cap
Bear Stun	Bear stuns hold for longer.	+0.2 s control; level 1 adds Bear Summon	35g	×1.55	5
Evolution Rate	Each level evolves the tower's damage output further.	+0.72 damage; level 1 adds Adaptive Evolution	45g	×1.65	no cap
Pack Bond	All summon auras buff allies more strongly.	+1.2% aura strength	30g	×1.5	no cap

Its schoolDruidism→Path AOvergrowth→Path BDruid→

StasisChronomancy path A, starting with Time Dilation and Temporal Decay.

School: ChronomancyPath: ARoles: crowd control · area damage · debuff

Abilities

On the tower from the moment it is built, before any upgrade.

Time Dilation

Time field slows enemies in range by 40% for 5 seconds.

Temporal Decay

Ages the target — ~1% max HP per second decay damage for 6 seconds.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Dilation Strength	Time-dilation field slows enemies inside harder.	+2% slow	25g	×1.5	no cap
Decay Stacks	Temporal-decay stack cap rises — compound stat-loss on the target.	+1 to the ability	30g	×1.55	10
Decay Strength	Each temporal-decay stack chews more %maxHP per second from the target.	+0.5 to the ability	28g	×1.5	no cap
Time Lock Count	Time-lock catches more enemies at once.	+1.5 damage; level 1 adds Time Lock	50g	×1.75	3
Lock Duration	Time-lock holds targets for longer.	+0.5 s control	40g	×1.6	6
Age to Dust	Age-to-Dust execute triggers at higher stack counts.	+0.5 damage; level 1 adds Age to Dust	45g	×1.65	no cap
Field Radius	Dilation field covers more cells.	+0.2 splash radius	30g	×1.5	8

Its schoolChronomancy→Path BRewind→Path CHaste→

RewindChronomancy path B, starting with Temporal Rewind.

School: ChronomancyPath: BRoles: support · crowd control · single-target damage

Abilities

On the tower from the moment it is built, before any upgrade.

Temporal Rewind

Rewinds the target along the path by ~2 tiles.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Rewind Distance	Rewind shoves enemies back further along their path.	+0.5 knockback	25g	×1.5	no cap
Chrono Boost	Allied towers in aura fire faster.	+2% aura strength; level 1 adds Chrono Boost	30g	×1.55	no cap
Loop Duration	Rewind loop shoves targets back further.	+0.3 to the ability; level 1 adds Time Loop	45g	×1.65	5
Loop Repeats	Rewind loop repeats stack more knockback (with Loop Duration).	+0.5 to the ability	50g	×1.75	3
Echo Strength	Echo-shots deal a higher % of the tower's hit damage.	+5 to the ability; level 1 adds Temporal Echo	35g	×1.55	no cap
Echo Frequency	Echo-shots fire more often (interval shrinks, floor 0.2s).	+0.1 to the ability	40g	×1.6	6
Rewind Frequency	Tower fires faster — rewinds trigger more often.	+0.05 shots/s	30g	×1.5	no cap

Its schoolChronomancy→Path AStasis→Path CHaste→

HasteChronomancy path C, starting with Aura Attack Speed 15 and Aging Curse.

School: ChronomancyPath: CRoles: support · area damage · damage over time

Abilities

On the tower from the moment it is built, before any upgrade.

Aura Attack Speed 15

Carried by every tower on this path.

Aging Curse

Curse of age: ~0.8% max HP per second over 5 seconds.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Aura AS	Allied towers in haste aura fire faster.	+2% aura strength	25g	×1.5	no cap
Aging Damage	Aging-curse DoT chews more % maxHP per second.	+0.1 damage	28g	×1.55	no cap
Confusion Chance	Higher roll that confusion seizes the target — briefly locks them in place.	+1.5 to the ability; level 1 adds Confusion	30g	×1.5	8
Entropy Slow	Entropy field slows enemies inside harder.	+2% slow; level 1 adds Entropy Field	25g	×1.5	no cap
Entropy DoT	Each entropy stack chews more % maxHP per second.	+0.1 damage	35g	×1.6	no cap
Epoch Duration	Epoch-burst stun and CC effects hold for longer.	+0.25 s control; level 1 adds Epoch Burst	45g	×1.65	5
Epoch Multiplier	Each level multiplies the tower's hit damage further.	+1 damage	40g	×1.6	6

Its schoolChronomancy→Path AStasis→Path BRewind→

SwarmBeastmastery path A, starting with Insect Swarm.

School: BeastmasteryPath: ARoles: area damage · damage over time · crowd control

Abilities

On the tower from the moment it is built, before any upgrade.

Insect Swarm

Releases swarms of insects that poison every nearby enemy for 5s.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Swarm Size	Swarm DoT ticks harder per level (bigger swarm = more bites).	+0.2 to the ability	25g	×1.5	no cap
Swarm Damage	Swarm DoT ticks harder on each target.	+0.25 damage	22g	×1.5	no cap
Split Count	Swarm splits to more enemies on a kill.	+1.5 damage; level 1 adds Swarm Split	40g	×1.65	3
Swarm Slow	Swarm slows targets harder while attached.	+2% slow	20g	×1.45	10
Armor Shred	Swarm strips more armor on every tick.	+0.3 damage	28g	×1.5	no cap
Spawn Rate	Swarm spawns more often (tick interval shrinks, floor 0.2s).	+0.05 to the ability	30g	×1.55	6
Devouring Scale	Devouring plague ticks with extra raw damage per level.	+2 to the ability; level 1 adds Devouring Plague	35g	×1.55	no cap

Its schoolBeastmastery→Path BAlpha→Path CChimera→

AlphaBeastmastery path B, starting with Alpha Beast.

School: BeastmasteryPath: BRoles: single-target damage · crowd control · support

Abilities

On the tower from the moment it is built, before any upgrade.

Alpha Beast

Summons a powerful alpha beast that bites enemies for major damage.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Alpha Damage	Alpha bleed ticks harder on the target per level.	+0.2 to the ability	28g	×1.55	no cap
Alpha HP	Alpha bleed lingers on the target for longer.	+0.2 to the ability	25g	×1.5	no cap
Charge Stun	Beast charge stuns hold for longer.	+0.2 s control	35g	×1.55	6
Roar Fear	Higher roll that the alpha's roar fears nearby enemies.	+5% proc chance	30g	×1.5	8
Regen Rate	Alpha heals between rakes — each bleed tick lands harder (stacks with Alpha Damage).	+0.5 to the ability	22g	×1.45	6
Pack Leader	All summoned beasts deal more damage.	+2% aura strength; level 1 adds Pack Leader	40g	×1.6	no cap
Pack Rotation	Tower fires faster.	+0.1 shots/s	35g	×1.55	6

Its schoolBeastmastery→Path ASwarm→Path CChimera→

ChimeraBeastmastery path C, starting with Spawn Mutant.

School: BeastmasteryPath: CRoles: area damage · debuff · single-target damage

Abilities

On the tower from the moment it is built, before any upgrade.

Spawn Mutant

Spawns a mutant beast with a random elemental tinge each time.

Upgrade slots (10)

* Damage, Attack Speed and Range are on every path; the other 7 are this path's own. “Gain per level” is what one level changes on the tower as the game computes it — several effect upgrades (damage over time, crits, shred, extra targets) count as extra hit damage, and “to the ability” is the raw amount the path's own ability is scaled by. Each level costs the previous level's price times the cost factor, so breadth is cheaper than depth.

Slot	What it does	Gain per level	Level-1 cost	Cost × per level	Max level
Damage *	Each level increases the tower's raw hit damage.	+3 damage	15g	×1.5	15 (extra levels do nothing)
Attack Speed *	Tower fires faster — more shots land per second.	+0.12 shots/s	15g	×1.5	15 (extra levels do nothing)
Range *	Stretches the tower's targeting reach further out.	+0.3 range	18g	×1.45	6
Mutant Variety	Each mutant variant bleeds harder onto enemies.	+1 to the ability	30g	×1.55	6
Mutant Stats	Each level strengthens the tower's mutants further.	+0.9 damage	25g	×1.5	no cap
Fusion Bonus	Hybrid fusion shot deals more raw damage per level.	+10 to the ability; level 1 adds Hybrid Fusion	35g	×1.55	no cap
Spawn Rate	Mutants spawn more often (tick interval shrinks, floor 0.2s).	+0.1 to the ability	28g	×1.5	6
Wave Reading	Strips more armor on every hit (wave-reading scales the shred).	+0.8 to the ability; level 1 adds Wave Reading	40g	×1.6	8
Unstable Explosion	Mutants explode harder on death — wider AoE.	+0.2 splash radius; level 1 adds Unstable Mutation	30g	×1.5	6
Apex Chimera	Tower's pack-leader aura grows stronger per level.	+0.6% aura strength	50g	×1.7	3

Its schoolBeastmastery→Path ASwarm→Path BAlpha→